On the Pitfalls of Analyzing Individual Neurons in Language Models
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Abstract (arXiv)
While many studies have shown that linguistic information is encoded in hidden word representations, few have studied individual neurons, to show how and in which neurons it is encoded. Among these, the common approach is to use an external probe to rank neurons according to their relevance to some linguistic attribute, and to evaluate the obtained ranking using the same probe that produced it. We show two pitfalls in this methodology: 1. It confounds distinct factors: probe quality and ranking quality. We separate them and draw conclusions on each. 2. It focuses on encoded information, rather than information that is used by the model. We show that these are not the same. We compare two recent ranking methods and a simple one we introduce, and evaluate them with regard to both of these aspects.
Key facts. Cited by 61 papers in the map (data through 7 Oct 2026). Builds on 18 of its 27 references that are in the map. First posted on arXiv 14 Oct 2021.
Builds on
18 of 27 references.
References · 18
- 2018BERT: Pre-training of Deep Bidirectional Transformers for Language Understandingcited by 40,204
- 2019Unsupervised Cross-lingual Representation Learning at Scalecited by 3,907
- 2019Analyzing Multi-Head Self-Attention: Specialized Heads Do the Heavy Lifting, the Rest Can Be Prunedcited by 844
- 2019Designing and Interpreting Probes with Control Taskscited by 608
- 2018What you can cram into a single vector: Probing sentence embeddings for linguistic propertiescited by 452
- 2016Fine-grained Analysis of Sentence Embeddings Using Auxiliary Prediction Taskscited by 216
- 2018What Is One Grain of Sand in the Desert? Analyzing Individual Neurons in Deep NLP Modelscited by 124
- 2020CausaLM: Causal Model Explanation Through Counterfactual Language Modelscited by 117
- 2018Identifying and Controlling Important Neurons in Neural Machine Translationcited by 110
- 2018On the importance of single directions for generalizationcited by 99
- 2018Under the Hood: Using Diagnostic Classifiers to Investigate and Improve how Language Models Track Agreement Informationcited by 97
- 2020Analyzing Individual Neurons in Pre-trained Language Modelscited by 97
6 more
- 2019The emergence of number and syntax units in LSTM language modelscited by 92
- 2021Counterfactual Interventions Reveal the Causal Effect of Relative Clause Representations on Agreement Predictioncited by 58
- 2020Intrinsic Probing through Dimension Selectioncited by 45
- 2018UniMorph 2.0: Universal Morphologycited by 40
- 2020It's not Greek to mBERT: Inducing Word-Level Translations from Multilingual BERTcited by 30
- 2018Marrying Universal Dependencies and Universal Morphologycited by 11
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Citing papers per year: 2021: 2; 2022: 10; 2023: 11; 2024: 16; 2025: 14; 2026: 8.
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- 2023Explainability for Large Language Models: A Surveycited by 331
- 2023Finding Neurons in a Haystack: Case Studies with Sparse Probingcited by 328
- 2023Emergent Linear Representations in World Models of Self-Supervised Sequence Modelscited by 313
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- 2022Toward Transparent AI: A Survey on Interpreting the Inner Structures of Deep Neural Networkscited by 126
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- 2024A Practical Review of Mechanistic Interpretability for Transformer-Based Language Modelscited by 113
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- 2023A Mechanistic Interpretation of Arithmetic Reasoning in Language Models using Causal Mediation Analysiscited by 88
- 2022Finding Skill Neurons in Pre-trained Transformer-based Language Modelscited by 65
- 2022Measuring Causal Effects of Data Statistics on Language Model's `Factual' Predictionscited by 57
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- 2021Sparse Interventions in Language Models with Differentiable Maskingcited by 34
- 2025A Survey on Mechanistic Interpretability for Multi-Modal Foundation Modelscited by 34
- 2025Adaptive Tool Use in Large Language Models with Meta-Cognition Triggercited by 29
- 2024Large Language Models Relearn Removed Conceptscited by 26
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- 2024Decomposing and Editing Predictions by Modeling Model Computationcited by 19
- 2024BadActs: A Universal Backdoor Defense in the Activation Spacecited by 19
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- 2023Can LLMs facilitate interpretation of pre-trained language models?cited by 15
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- 2022Same Neurons, Different Languages: Probing Morphosyntax in Multilingual Pre-trained Modelscited by 14
- 2022Causal Analysis of Syntactic Agreement Neurons in Multilingual Language Modelscited by 14
- 2024Tracing the Roots of Facts in Multilingual Language Models: Independent, Shared, and Transferred Knowledgecited by 14
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- 2021Probing as Quantifying Inductive Biascited by 11
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- 2025Neurons Speak in Ranges: Breaking Free from Discrete Neuronal Attributioncited by 5
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- 2025Distilling Knowledge from Large Language Models: A Concept Bottleneck Model for Hate and Counter Speech Recognitioncited by 4
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- 2025Disentangling Linguistic Features with Dimension-Wise Analysis of Vector Embeddingscited by 2
50 most cited of 61 All 61 (JSON)
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- 2026Representation Measurements Under Function-Preserving Reparameterizations27 Aug 2026
- 2026Towards Effective In-context Cross-domain Knowledge Transfer via Domain-invariant-neurons-based Retrieval7 Apr 2026
- 2026NeST: Neuron Selective Tuning for LLM Safety18 Feb 2026
- 2026A Study of Crosslinguistic Influence in Language Models29 Jan 2026
- 2026Discovering and Causally Validating Emotion-Sensitive Neurons in Large Audio-Language Models6 Jan 2026
- 2025Unboxing the Black Box: Mechanistic Interpretability for Algorithmic Understanding of Neural Networks24 Nov 2025
- 2025Equivariant Sparse Autoencoders: Mechanistic Interpretability of Neural Networks on Symmetric Data12 Nov 2025
- 2025Continual Personalization for Diffusion Models2 Oct 2025
- 2025Do Activation Verbalization Methods Convey Privileged Information?16 Sep 2025
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- 2025Distilling Knowledge from Large Language Models: A Concept Bottleneck Model for Hate and Counter Speech Recognition30 Jul 2025
- 2025Cross-Layer Discrete Concept Discovery for Interpreting Language Models24 Jun 2025
- 2025Disentangling Linguistic Features with Dimension-Wise Analysis of Vector Embeddings20 Apr 2025
- 2025On Relation-Specific Neurons in Large Language Models24 Feb 2025
- 2025A Survey on Mechanistic Interpretability for Multi-Modal Foundation Models22 Feb 2025
- 2025Adaptive Tool Use in Large Language Models with Meta-Cognition Trigger18 Feb 2025
- 2025Neurons Speak in Ranges: Breaking Free from Discrete Neuronal Attribution4 Feb 2025
- 2025Enhancing Automated Interpretability with Output-Centric Feature Descriptions14 Jan 2025
- 2024A Theoretical Survey on Foundation Models15 Oct 2024
- 2024Mechanistic?7 Oct 2024
- 2024The Quest for the Right Mediator: Surveying Mechanistic Interpretability Through the Lens of Causal Mediation Analysis2 Aug 2024
- 2024On Behalf of the Stakeholders: Trends in NLP Model Interpretability in the Era of LLMs27 Jul 2024
- 2024Crafting Large Language Models for Enhanced Interpretability5 Jul 2024
- 2024A Practical Review of Mechanistic Interpretability for Transformer-Based Language Models2 Jul 2024
- 2024Confidence Regulation Neurons in Language Models24 Jun 2024
- 2024ConceptPrune: Concept Editing in Diffusion Models via Skilled Neuron Pruning29 May 2024
- 2024BadActs: A Universal Backdoor Defense in the Activation Space18 May 2024
- 2024Decomposing and Editing Predictions by Modeling Model Computation17 Apr 2024
- 2024Towards detecting unanticipated bias in Large Language Models3 Apr 2024
- 2024Tracing the Roots of Facts in Multilingual Language Models: Independent, Shared, and Transferred Knowledge8 Mar 2024
- 2024Assessing the Brittleness of Safety Alignment via Pruning and Low-Rank Modifications7 Feb 2024
- 2024Black-Box Access is Insufficient for Rigorous AI Audits25 Jan 2024
- 2024Universal Neurons in GPT2 Language Models22 Jan 2024
- 2024Large Language Models Relearn Removed Concepts3 Jan 2024
- 2023Towards a fuller understanding of neurons with Clustered Compositional Explanations27 Oct 2023
- 2023DeepDecipher: Accessing and Investigating Neuron Activation in Large Language Models3 Oct 2023
- 2023Rigorously Assessing Natural Language Explanations of Neurons19 Sep 2023
- 2023Explainability for Large Language Models: A Survey2 Sep 2023
- 2023Emergent Linear Representations in World Models of Self-Supervised Sequence Models2 Sep 2023
- 2023NeuroX Library for Neuron Analysis of Deep NLP Models26 May 2023
- 2023A Mechanistic Interpretation of Arithmetic Reasoning in Language Models using Causal Mediation Analysis24 May 2023
- 2023Can LLMs facilitate interpretation of pre-trained language models?22 May 2023
- 2023Finding Neurons in a Haystack: Case Studies with Sparse Probing2 May 2023
- 2023Redundancy and Concept Analysis for Code-trained Language Models1 May 2023
- 2023Evaluating Neuron Interpretation Methods of NLP Models30 Jan 2023
- 2022Finding Skill Neurons in Pre-trained Transformer-based Language Models14 Nov 2022
50 newest of 61 All 61 (JSON)
- 2024Assessing the Brittleness of Safety Alignment via Pruning and Low-Rank Modificationscited by 220
- 2025NeuroStrike: Neuron-Level Attacks on Aligned LLMscited by 18
- 2026NeST: Neuron Selective Tuning for LLM Safetycited by 2
- 2023Explainability for Large Language Models: A Surveycited by 331
- 2023Finding Neurons in a Haystack: Case Studies with Sparse Probingcited by 328
- 2022Toward Transparent AI: A Survey on Interpreting the Inner Structures of Deep Neural Networkscited by 126
- 2024Black-Box Access is Insufficient for Rigorous AI Auditscited by 123
- 2024A Practical Review of Mechanistic Interpretability for Transformer-Based Language Modelscited by 113
- 2024Universal Neurons in GPT2 Language Modelscited by 100
- 2022Finding Skill Neurons in Pre-trained Transformer-based Language Modelscited by 65
- 2022Measuring Causal Effects of Data Statistics on Language Model's `Factual' Predictionscited by 57
- 2024Confidence Regulation Neurons in Language Modelscited by 57
- 2023Rigorously Assessing Natural Language Explanations of Neuronscited by 44
- 2025Enhancing Automated Interpretability with Output-Centric Feature Descriptionscited by 39
- 2021Sparse Interventions in Language Models with Differentiable Maskingcited by 34
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- 2026KDFP: A first-principles approach to knowledge distillation in large language models7 Oct 2026
- 2026Grammatical "grandmother neurons" are rare in LLMs24 Sep 2026
- 2026A Mechanistic Study of AI-Text Detection Neurons in Frozen BERT: Sparse Probing and Activation Patching on RAID8 Sep 2026
- 2026Representation Measurements Under Function-Preserving Reparameterizations27 Aug 2026
- 2026A Study of Crosslinguistic Influence in Language Models29 Jan 2026
- 2026Discovering and Causally Validating Emotion-Sensitive Neurons in Large Audio-Language Models6 Jan 2026
- 2025Unboxing the Black Box: Mechanistic Interpretability for Algorithmic Understanding of Neural Networks24 Nov 2025
- 2025Equivariant Sparse Autoencoders: Mechanistic Interpretability of Neural Networks on Symmetric Data12 Nov 2025
- 2025Do Activation Verbalization Methods Convey Privileged Information?16 Sep 2025
- 2025Evaluating Sparse Autoencoders for Monosemantic Representation20 Aug 2025
- 2025Distilling Knowledge from Large Language Models: A Concept Bottleneck Model for Hate and Counter Speech Recognition30 Jul 2025
- 2025Cross-Layer Discrete Concept Discovery for Interpreting Language Models24 Jun 2025
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