What Is One Grain of Sand in the Desert? Analyzing Individual Neurons in Deep NLP Models
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Abstract (arXiv)
Despite the remarkable evolution of deep neural networks in natural language processing (NLP), their interpretability remains a challenge. Previous work largely focused on what these models learn at the representation level. We break this analysis down further and study individual dimensions (neurons) in the vector representation learned by end-to-end neural models in NLP tasks. We propose two methods: Linguistic Correlation Analysis, based on a supervised method to extract the most relevant neurons with respect to an extrinsic task, and Cross-model Correlation Analysis, an unsupervised method to extract salient neurons w.r.t. the model itself. We evaluate the effectiveness of our techniques by ablating the identified neurons and reevaluating the network's performance for two tasks: neural machine translation (NMT) and neural language modeling (NLM). We further present a comprehensive analysis of neurons with the aim to address the following questions: i) how localized or distributed are different linguistic properties in the models? ii) are certain neurons exclusive to some properties and not others? iii) is the information more or less distributed in NMT vs. NLM? and iv) how important are the neurons identified through the linguistic correlation method to the overall task? Our code is publicly available as part of the NeuroX toolkit (Dalvi et al. 2019).
Key facts. Cited by 124 papers in the map (data through 7 Oct 2026). First posted on arXiv 21 Dec 2018.
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Citing papers per year: 2021: 17; 2022: 25; 2023: 17; 2024: 31; 2025: 24; 2026: 10.
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50 most cited of 124 All 124 (JSON)
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- 2024On the Universal Truthfulness Hyperplane Inside LLMs11 Jul 2024
- 2024Crafting Large Language Models for Enhanced Interpretability5 Jul 2024
- 2024A Practical Review of Mechanistic Interpretability for Transformer-Based Language Models2 Jul 2024
- 2024Confidence Regulation Neurons in Language Models24 Jun 2024
- 2024Identifying Query-Relevant Neurons in Large Language Models for Long-Form Texts16 Jun 2024
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- 2024A Multimodal Automated Interpretability Agentcited by 62
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- 2023The Geometry of Truth: Emergent Linear Structure in Large Language Model Representations of True/False Datasetscited by 673
- 2023Explainability for Large Language Models: A Surveycited by 331
- 2023Finding Neurons in a Haystack: Case Studies with Sparse Probingcited by 328
- 2023How does GPT-2 compute greater-than?: Interpreting mathematical abilities in a pre-trained language modelcited by 199
- 2022Natural Language Descriptions of Deep Visual Featurescited by 136
- 2022Analyzing Transformers in Embedding Spacecited by 131
- 2024Language-Specific Neurons: The Key to Multilingual Capabilities in Large Language Modelscited by 124
- 2022The Lazy Neuron Phenomenon: On Emergence of Activation Sparsity in Transformerscited by 121
- 2024A Practical Review of Mechanistic Interpretability for Transformer-Based Language Modelscited by 113
- 2024Universal Neurons in GPT2 Language Modelscited by 100
- 2021An Interpretability Illusion for BERTcited by 94
- 2022Head2Toe: Utilizing Intermediate Representations for Better Transfer Learningcited by 83
12 most cited of 114 All 114 (JSON)
- 2026Grammatical "grandmother neurons" are rare in LLMs24 Sep 2026
- 2026A Mechanistic Study of AI-Text Detection Neurons in Frozen BERT: Sparse Probing and Activation Patching on RAID8 Sep 2026
- 2026Emotion Across Speech and Faces: Shared Affective Mechanisms in Multimodal Foundation Models17 Aug 2026
- 2026Multilingual Emotion Neurons in Large Audio-Language Models9 Aug 2026
- 2026Explaining Attention with Program Synthesis17 Jun 2026
- 2026Neuron-Level Emotion Control in Speech-Generative Large Audio-Language Models18 Mar 2026
- 2026SYNAPSE: Framework for Neuron Analysis and Perturbation in Sequence Encoding9 Mar 2026
- 2026GLUScope: A Tool for Analyzing GLU Neurons in Transformer Language Models27 Feb 2026
- 2026CRANE: Causal Relevance Analysis of Language-Specific Neurons in Multilingual Large Language Models8 Jan 2026
- 2026Discovering and Causally Validating Emotion-Sensitive Neurons in Large Audio-Language Models6 Jan 2026
- 2025One Swallow Does Not Make a Summer: Understanding Semantic Structures in Embedding Spaces30 Nov 2025
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