Anatomy of a Machine Learning Ecosystem: 2 Million Models on Hugging Face
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Abstract (arXiv)
Many have observed that the development and deployment of generative machine learning (ML) and artificial intelligence (AI) models follow a distinctive pattern in which pre-trained models are adapted and fine-tuned for specific downstream tasks. However, there is limited empirical work that examines the structure of these interactions. This paper analyzes 1.86 million models on Hugging Face, a leading peer production platform for model development. Our study of model family trees -- networks that connect fine-tuned models to their base or parent -- reveals sprawling fine-tuning lineages that vary widely in size and structure. Using an evolutionary biology lens to study ML models, we use model metadata and model cards to measure the genetic similarity and mutation of traits over model families. We find that models tend to exhibit a family resemblance, meaning their genetic markers and traits exhibit more overlap when they belong to the same model family. However, these similarities depart in certain ways from standard models of asexual reproduction, because mutations are fast and directed, such that two `sibling' models tend to exhibit more similarity than parent/child pairs. Further analysis of the directional drifts of these mutations reveals qualitative insights about the open machine learning ecosystem: Licenses counter-intuitively drift from restrictive, commercial licenses towards permissive or copyleft licenses, often in violation of upstream license's terms; models evolve from multi-lingual compatibility towards english-only compatibility; and model cards reduce in length and standardize by turning, more often, to templates and automatically generated text. Overall, this work takes a step toward an empirically grounded understanding of model fine-tuning and suggests that ecological models and methods can yield novel scientific insights.
Key facts. Cited by 12 papers in the map (data through 14 Oct 2026). Builds on 25 of its 54 references that are in the map. First posted on arXiv 9 Aug 2025.
Builds on
25 of 54 references.
References · 25
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13 more
- 2020A Survey on Contextual Embeddingscited by 48
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Built on by
Citing papers per year: 2025: 2; 2026: 10.
12 papers.
- 2026Efficient Evaluation of LLM Performance with Statistical Guaranteescited by 18
- 2025Economies of Open Intelligence: Tracing Power & Participation in the Model Ecosystemcited by 4
- 2026EVOMAL: Self-Poisoning in Self-Evolving Coding Agentscited by 3
- 2025A Cartography of Open Collaboration in Open Source AI: Mapping Practices, Motivations, and Governance in 14 Open Large Language Model Projectscited by 1
- 2026Which Models Are Our Models Built On? Auditing Invisible Dependencies in Modern LLMscited by 1
- 2026AdaQE-CG: Adaptive Query Expansion for Web-Scale Generative AI Model and Data Card Generationcited by 0
- 2026TensorDex: A Compact, Tensor-Centric Storage System for Modern AI Modelscited by 0
- 2026Cost-Ordered Feasibility for Multi-Armed Bandits with Cost Subsidycited by 0
- 2026ArtifactLinker: Linking Scientific Artifacts for Automatic State-of-the-Art Discoverycited by 0
- 2026A governance horizon for ethical-use constraints in open-weight AI modelscited by 0
- 2026AI Supply Chain Galaxy: 3D Visual Analytics for License Compliancecited by 0
- 2026The evolution of sex for artificial intelligence: a population-genetic framework for multigenerational model populationscited by 0
- 2026The evolution of sex for artificial intelligence: a population-genetic framework for multigenerational model populations16 Sep 2026
- 2026EVOMAL: Self-Poisoning in Self-Evolving Coding Agents26 Aug 2026
- 2026AI Supply Chain Galaxy: 3D Visual Analytics for License Compliance15 Jun 2026
- 2026Which Models Are Our Models Built On? Auditing Invisible Dependencies in Modern LLMs10 Jun 2026
- 2026A governance horizon for ethical-use constraints in open-weight AI models23 May 2026
- 2026ArtifactLinker: Linking Scientific Artifacts for Automatic State-of-the-Art Discovery16 May 2026
- 2026Cost-Ordered Feasibility for Multi-Armed Bandits with Cost Subsidy8 May 2026
- 2026TensorDex: A Compact, Tensor-Centric Storage System for Modern AI Models18 Apr 2026
- 2026AdaQE-CG: Adaptive Query Expansion for Web-Scale Generative AI Model and Data Card Generation16 Mar 2026
- 2026Efficient Evaluation of LLM Performance with Statistical Guarantees28 Jan 2026
- 2025Economies of Open Intelligence: Tracing Power & Participation in the Model Ecosystem27 Nov 2025
- 2025A Cartography of Open Collaboration in Open Source AI: Mapping Practices, Motivations, and Governance in 14 Open Large Language Model Projects29 Sep 2025