What does the Knowledge Neuron Thesis Have to do with Knowledge?
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Abstract (arXiv)
We reassess the Knowledge Neuron (KN) Thesis: an interpretation of the mechanism underlying the ability of large language models to recall facts from a training corpus. This nascent thesis proposes that facts are recalled from the training corpus through the MLP weights in a manner resembling key-value memory, implying in effect that "knowledge" is stored in the network. Furthermore, by modifying the MLP modules, one can control the language model's generation of factual information. The plausibility of the KN thesis has been demonstrated by the success of KN-inspired model editing methods (Dai et al., 2022; Meng et al., 2022). We find that this thesis is, at best, an oversimplification. Not only have we found that we can edit the expression of certain linguistic phenomena using the same model editing methods but, through a more comprehensive evaluation, we have found that the KN thesis does not adequately explain the process of factual expression. While it is possible to argue that the MLP weights store complex patterns that are interpretable both syntactically and semantically, these patterns do not constitute "knowledge." To gain a more comprehensive understanding of the knowledge representation process, we must look beyond the MLP weights and explore recent models' complex layer structures and attention mechanisms.
Key facts. Cited by 49 papers in the map (data through 7 Oct 2026). Builds on 24 of its 32 references that are in the map. First posted on arXiv 3 May 2024.
Builds on
24 of 32 references.
References · 24
- 2018BERT: Pre-training of Deep Bidirectional Transformers for Language Understandingcited by 40,204
- 2019XLNet: Generalized Autoregressive Pretraining for Language Understandingcited by 2,801
- 2022Locating and Editing Factual Associations in GPTcited by 2,609
- 2019Language Models as Knowledge Bases?cited by 1,922
- 2020Transformer Feed-Forward Layers Are Key-Value Memoriescited by 1,307
- 2022Interpretability in the Wild: a Circuit for Indirect Object Identification in GPT-2 smallcited by 1,130
- 2019BERT Rediscovers the Classical NLP Pipelinecited by 1,046
- 2019What Does BERT Look At? An Analysis of BERT's Attentioncited by 994
- 2022Mass-Editing Memory in a Transformercited by 976
- 2023Towards Automated Circuit Discovery for Mechanistic Interpretabilitycited by 700
- 2019Designing and Interpreting Probes with Control Taskscited by 608
- 2021Knowledge Neurons in Pretrained Transformerscited by 590
12 more
- 2019BLiMP: The Benchmark of Linguistic Minimal Pairs for Englishcited by 553
- 2021Editing Factual Knowledge in Language Modelscited by 548
- 2021Fast Model Editing at Scalecited by 491
- 2017Zero-Shot Relation Extraction via Reading Comprehensioncited by 425
- 2023Editing Large Language Models: Problems, Methods, and Opportunitiescited by 367
- 2021Measuring and Improving Consistency in Pretrained Language Modelscited by 361
- 2023Does Localization Inform Editing? Surprising Differences in Causality-Based Localization vs. Knowledge Editing in Language Modelscited by 237
- 2021Causal Analysis of Syntactic Agreement Mechanisms in Neural Language Modelscited by 127
- 2023Interpretability at Scale: Identifying Causal Mechanisms in Alpacacited by 102
- 2020Analyzing Individual Neurons in Pre-trained Language Modelscited by 97
- 2023Rigorously Assessing Natural Language Explanations of Neuronscited by 44
- 2022Characterizing Intrinsic Compositionality in Transformers with Tree Projectionscited by 40
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Citing papers per year: 2024: 18; 2025: 17; 2026: 14.
49 papers.
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- 2024Transformers to SSMs: Distilling Quadratic Knowledge to Subquadratic Modelscited by 66
- 2024Knowledge Circuits in Pretrained Transformerscited by 60
- 2024NLSR: Neuron-Level Safety Realignment of Large Language Models Against Harmful Fine-Tuningcited by 37
- 2024Can Knowledge Editing Really Correct Hallucinations?cited by 28
- 2024MMNeuron: Discovering Neuron-Level Domain-Specific Interpretation in Multimodal Large Language Modelcited by 26
- 2024Style-Specific Neurons for Steering LLMs in Text Style Transfercited by 24
- 2024Can Editing LLMs Inject Harm?cited by 22
- 2024LLM-Generated Black-box Explanations Can Be Adversarially Helpfulcited by 20
- 2024Mitigating the Language Mismatch and Repetition Issues in LLM-based Machine Translation via Model Editingcited by 18
- 2024Has this Fact been Edited? Detecting Knowledge Edits in Language Modelscited by 14
- 2025Does Time Have Its Place? Temporal Heads: Where Language Models Recall Time-specific Informationcited by 14
- 2024Knowledge Localization: Mission Not Accomplished? Enter Query Localization!cited by 13
37 more
- 2024Sheaf Discovery with Joint Computation Graph Pruning and Flexible Granularitycited by 11
- 2025UORA: Uniform Orthogonal Reinitialization Adaptation in Parameter-Efficient Fine-Tuning of Large Modelscited by 11
- 2024The Geometry of Numerical Reasoning: Language Models Compare Numeric Properties in Linear Subspacescited by 10
- 2024Natural Language Processing RELIES on Linguisticscited by 9
- 2024Generalisation First, Memorisation Second? Memorisation Localisation for Natural Language Classification Taskscited by 9
- 2024Context-Parametric Inversion: Why Instruction Finetuning Can Worsen Context Reliancecited by 9
- 2026All Circuits Lead to Rome: Rethinking Functional Anisotropy in Circuit and Sheaf Discovery for LLMscited by 8
- 2025Llama See, Llama Do: A Mechanistic Perspective on Contextual Entrainment and Distraction in LLMscited by 6
- 2025COMPKE: Complex Question Answering under Knowledge Editingcited by 6
- 2025The Knowledge Microscope: Features as Better Analytical Lenses than Neuronscited by 5
- 2024How Well Can Knowledge Edit Methods Edit Perplexing Knowledge?cited by 4
- 2025Brain-Inspired Exploration of Functional Networks and Key Neurons in Large Language Modelscited by 4
- 2025Drop Dropout on Single-Epoch Language Model Pretrainingcited by 3
- 2025Spelling-out is not Straightforward: LLMs' Capability of Tokenization from Token to Characterscited by 3
- 2025Fine-Tuned LLMs Know They Don't Know: A Parameter-Efficient Approach to Recovering Honestycited by 3
- 2026Auditing Language Model Unlearning via Information Decompositioncited by 3
- 2026SHIFT: Gate-Modulated Activation Steering for Knowledge Conflict Mitigation in Retrieval-Augmented Generationcited by 3
- 2024Understanding Language Model Circuits through Knowledge Editingcited by 2
- 2025Truth Neuronscited by 2
- 2025Explainability of Large Language Models: Opportunities and Challenges toward Generating Trustworthy Explanationscited by 2
- 2026Attention Editing: A Versatile Framework for Cross-Architecture Attention Conversioncited by 2
- 2025Understanding Fact Recall in Language Models: Why Two-Stage Training Encourages Memorization but Mixed Training Teaches Knowledgecited by 1
- 2025Understanding Gated Neurons in Transformers from Their Input-Output Functionalitycited by 1
- 2025On the Representations of Entities in Auto-regressive Large Language Modelscited by 1
- 2025From Memorization to Generalization: Fine-Tuning Large Language Models for Biomedical Term-to-Identifier Normalizationcited by 1
- 2025SurvSurf: a partially monotonic neural network for first-hitting time prediction of intermittently observed discrete and continuous sequential eventscited by 0
- 2025Tracing Multilingual Knowledge Acquisition Dynamics in Domain Adaptation: A Case Study of English-Japanese Biomedical Adaptationcited by 0
- 2026Knowing Bias, Doing Better: Mitigating Social Bias in LLMs via Know-Bias Neuron Enhancementcited by 0
- 2026Consensus-Aligned Neuron Efficient Fine-Tuning Large Language Models for Multi-Domain Machine Translationcited by 0
- 2026GLUScope: A Tool for Analyzing GLU Neurons in Transformer Language Modelscited by 0
- 2026From Insight to Action: A Novel Framework for Interpretability-Guided Data Selection in Large Language Modelscited by 0
- 2026GKnow: Measuring the Entanglement of Gender Bias and Factual Gendercited by 0
- 2026RepTran: Search-Based Repair of Transformer Modelscited by 0
- 2026RACE: Scalable Statistical Estimation of Functional Consistency in LLM Neuronscited by 0
- 2026Some Tokens Behave like Magnets: Revealing Linguistic Organization in the Layers of Language Modelscited by 0
- 2026Weakening Neurons: An Input-Output Functionality in Transformers with Outsize Influencecited by 0
- 2026Hallucination Neurons and Where to Find Them: An Investigation into the existence of Hallucination Neuronscited by 0
- 2026Hallucination Neurons and Where to Find Them: An Investigation into the existence of Hallucination Neurons24 Sep 2026
- 2026Weakening Neurons: An Input-Output Functionality in Transformers with Outsize Influence16 Sep 2026
- 2026Some Tokens Behave like Magnets: Revealing Linguistic Organization in the Layers of Language Models4 Sep 2026
- 2026RACE: Scalable Statistical Estimation of Functional Consistency in LLM Neurons25 Aug 2026
- 2026RepTran: Search-Based Repair of Transformer Models13 Jul 2026
- 2026SHIFT: Gate-Modulated Activation Steering for Knowledge Conflict Mitigation in Retrieval-Augmented Generation26 Jun 2026
- 2026All Circuits Lead to Rome: Rethinking Functional Anisotropy in Circuit and Sheaf Discovery for LLMs12 May 2026
- 2026GKnow: Measuring the Entanglement of Gender Bias and Factual Gender12 May 2026
- 2026From Insight to Action: A Novel Framework for Interpretability-Guided Data Selection in Large Language Models28 Apr 2026
- 2026Attention Editing: A Versatile Framework for Cross-Architecture Attention Conversion7 Apr 2026
- 2026GLUScope: A Tool for Analyzing GLU Neurons in Transformer Language Models27 Feb 2026
- 2026Consensus-Aligned Neuron Efficient Fine-Tuning Large Language Models for Multi-Domain Machine Translation5 Feb 2026
37 more
- 2026Knowing Bias, Doing Better: Mitigating Social Bias in LLMs via Know-Bias Neuron Enhancement29 Jan 2026
- 2026Auditing Language Model Unlearning via Information Decomposition21 Jan 2026
- 2025Fine-Tuned LLMs Know They Don't Know: A Parameter-Efficient Approach to Recovering Honesty17 Nov 2025
- 2025From Memorization to Generalization: Fine-Tuning Large Language Models for Biomedical Term-to-Identifier Normalization21 Oct 2025
- 2025Explainability of Large Language Models: Opportunities and Challenges toward Generating Trustworthy Explanations20 Oct 2025
- 2025Tracing Multilingual Knowledge Acquisition Dynamics in Domain Adaptation: A Case Study of English-Japanese Biomedical Adaptation14 Oct 2025
- 2025On the Representations of Entities in Auto-regressive Large Language Models10 Oct 2025
- 2025Spelling-out is not Straightforward: LLMs' Capability of Tokenization from Token to Characters12 Jun 2025
- 2025COMPKE: Complex Question Answering under Knowledge Editing1 Jun 2025
- 2025Drop Dropout on Single-Epoch Language Model Pretraining30 May 2025
- 2025UORA: Uniform Orthogonal Reinitialization Adaptation in Parameter-Efficient Fine-Tuning of Large Models26 May 2025
- 2025Understanding Gated Neurons in Transformers from Their Input-Output Functionality23 May 2025
- 2025Understanding Fact Recall in Language Models: Why Two-Stage Training Encourages Memorization but Mixed Training Teaches Knowledge22 May 2025
- 2025Truth Neurons18 May 2025
- 2025Llama See, Llama Do: A Mechanistic Perspective on Contextual Entrainment and Distraction in LLMs14 May 2025
- 2025SurvSurf: a partially monotonic neural network for first-hitting time prediction of intermittently observed discrete and continuous sequential events7 Apr 2025
- 2025Does Time Have Its Place? Temporal Heads: Where Language Models Recall Time-specific Information20 Feb 2025
- 2025The Knowledge Microscope: Features as Better Analytical Lenses than Neurons18 Feb 2025
- 2025Brain-Inspired Exploration of Functional Networks and Key Neurons in Large Language Models13 Feb 2025
- 2024NLSR: Neuron-Level Safety Realignment of Large Language Models Against Harmful Fine-Tuning17 Dec 2024
- 2024Can Knowledge Editing Really Correct Hallucinations?21 Oct 2024
- 2024The Geometry of Numerical Reasoning: Language Models Compare Numeric Properties in Linear Subspaces17 Oct 2024
- 2024Context-Parametric Inversion: Why Instruction Finetuning Can Worsen Context Reliance14 Oct 2024
- 2024Mitigating the Language Mismatch and Repetition Issues in LLM-based Machine Translation via Model Editing9 Oct 2024
- 2024Style-Specific Neurons for Steering LLMs in Text Style Transfer1 Oct 2024
- 2024Transformers to SSMs: Distilling Quadratic Knowledge to Subquadratic Models19 Aug 2024
- 2024Generalisation First, Memorisation Second? Memorisation Localisation for Natural Language Classification Tasks9 Aug 2024
- 2024Can Editing LLMs Inject Harm?29 Jul 2024
- 2024Sheaf Discovery with Joint Computation Graph Pruning and Flexible Granularity4 Jul 2024
- 2024How Well Can Knowledge Edit Methods Edit Perplexing Knowledge?25 Jun 2024
- 2024Understanding Language Model Circuits through Knowledge Editing25 Jun 2024
- 2024MMNeuron: Discovering Neuron-Level Domain-Specific Interpretation in Multimodal Large Language Model17 Jun 2024
- 2024Knowledge Circuits in Pretrained Transformers28 May 2024
- 2024Knowledge Localization: Mission Not Accomplished? Enter Query Localization!23 May 2024
- 2024LLM-Generated Black-box Explanations Can Be Adversarially Helpful10 May 2024
- 2024Natural Language Processing RELIES on Linguistics9 May 2024
- 2024Has this Fact been Edited? Detecting Knowledge Edits in Language Models4 May 2024
- 2024Can Editing LLMs Inject Harm?cited by 22
- 2026Auditing Language Model Unlearning via Information Decompositioncited by 3
- 2026Auditing Language Model Unlearning via Information Decomposition21 Jan 2026
- 2024Can Editing LLMs Inject Harm?29 Jul 2024
- 2024Knowledge Circuits in Pretrained Transformerscited by 60
- 2024Can Knowledge Editing Really Correct Hallucinations?cited by 28
- 2024Style-Specific Neurons for Steering LLMs in Text Style Transfercited by 24
- 2024LLM-Generated Black-box Explanations Can Be Adversarially Helpfulcited by 20
- 2024Mitigating the Language Mismatch and Repetition Issues in LLM-based Machine Translation via Model Editingcited by 18
- 2024Has this Fact been Edited? Detecting Knowledge Edits in Language Modelscited by 14
- 2025Does Time Have Its Place? Temporal Heads: Where Language Models Recall Time-specific Informationcited by 14
- 2024Knowledge Localization: Mission Not Accomplished? Enter Query Localization!cited by 13
- 2024Sheaf Discovery with Joint Computation Graph Pruning and Flexible Granularitycited by 11
- 2025UORA: Uniform Orthogonal Reinitialization Adaptation in Parameter-Efficient Fine-Tuning of Large Modelscited by 11
- 2024Natural Language Processing RELIES on Linguisticscited by 9
- 2024Generalisation First, Memorisation Second? Memorisation Localisation for Natural Language Classification Taskscited by 9
12 most cited of 40 All 40 (JSON)
- 2026Weakening Neurons: An Input-Output Functionality in Transformers with Outsize Influence16 Sep 2026
- 2026Some Tokens Behave like Magnets: Revealing Linguistic Organization in the Layers of Language Models4 Sep 2026
- 2026RACE: Scalable Statistical Estimation of Functional Consistency in LLM Neurons25 Aug 2026
- 2026RepTran: Search-Based Repair of Transformer Models13 Jul 2026
- 2026GKnow: Measuring the Entanglement of Gender Bias and Factual Gender12 May 2026
- 2026From Insight to Action: A Novel Framework for Interpretability-Guided Data Selection in Large Language Models28 Apr 2026
- 2026Attention Editing: A Versatile Framework for Cross-Architecture Attention Conversion7 Apr 2026
- 2026GLUScope: A Tool for Analyzing GLU Neurons in Transformer Language Models27 Feb 2026
- 2026Consensus-Aligned Neuron Efficient Fine-Tuning Large Language Models for Multi-Domain Machine Translation5 Feb 2026
- 2026Knowing Bias, Doing Better: Mitigating Social Bias in LLMs via Know-Bias Neuron Enhancement29 Jan 2026
- 2025Fine-Tuned LLMs Know They Don't Know: A Parameter-Efficient Approach to Recovering Honesty17 Nov 2025
- 2025From Memorization to Generalization: Fine-Tuning Large Language Models for Biomedical Term-to-Identifier Normalization21 Oct 2025
12 newest of 40 All 40 (JSON)