Probing artificial neural networks: insights from neuroscience
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Abstract (arXiv)
A major challenge in both neuroscience and machine learning is the development of useful tools for understanding complex information processing systems. One such tool is probes, i.e., supervised models that relate features of interest to activation patterns arising in biological or artificial neural networks. Neuroscience has paved the way in using such models through numerous studies conducted in recent decades. In this work, we draw insights from neuroscience to help guide probing research in machine learning. We highlight two important design choices for probes - direction and expressivity - and relate these choices to research goals. We argue that specific research goals play a paramount role when designing a probe and encourage future probing studies to be explicit in stating these goals.
Key facts. Cited by 17 papers in the map (data through 7 Oct 2026). Builds on 14 of its 42 references that are in the map. First posted on arXiv 16 Apr 2021.
Builds on
14 of 42 references.
References · 14
- 2016Understanding intermediate layers using linear classifier probescited by 1,279
- 2019Designing and Interpreting Probes with Control Taskscited by 608
- 2019Linguistic Knowledge and Transferability of Contextual Representationscited by 382
- 2020Information-Theoretic Probing with Minimum Description Lengthcited by 230
- 2016Fine-grained Analysis of Sentence Embeddings Using Auxiliary Prediction Taskscited by 216
- 2020Information-Theoretic Probing for Linguistic Structurecited by 204
- 2020Probing the Probing Paradigm: Does Probing Accuracy Entail Task Relevance?cited by 144
- 2018Under the Hood: Using Diagnostic Classifiers to Investigate and Improve how Language Models Track Agreement Informationcited by 97
- 2020Do Language Embeddings Capture Scales?cited by 63
- 2020Pareto Probing: Trading Off Accuracy for Complexitycited by 55
- 2021Neural population geometry: An approach for understanding biological and artificial neural networkscited by 52
- 2020Do Neural Language Models Show Preferences for Syntactic Formalisms?cited by 30
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Citing papers per year: 2021: 2; 2022: 1; 2023: 6; 2024: 3; 2025: 3; 2026: 2.
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- 2021Conditional probing: measuring usable information beyond a baselinecited by 61
- 2023SkinCon: A skin disease dataset densely annotated by domain experts for fine-grained model debugging and analysiscited by 44
- 2023Dissociating language and thought in large language modelscited by 27
- 2024Language Models as Models of Languagecited by 24
- 2025Representation Engineering for Large-Language Models: Survey and Research Challengescited by 18
- 2021Neural Population Geometry Reveals the Role of Stochasticity in Robust Perceptioncited by 12
- 2023Probing Graph Representationscited by 10
- 2026From Weights to Activations: Is Steering the Next Frontier of Adaptation?cited by 8
- 2022Trustworthy Social Bias Measurementcited by 7
- 2024Multilevel Interpretability Of Artificial Neural Networks: Leveraging Framework And Methods From Neurosciencecited by 5
- 2024Vision-Language Models Align with Human Neural Representations in Concept Processingcited by 2
- 2025Prophecy: Inferring Formal Properties from Neuron Activationscited by 2
5 more
- 2023Language Cognition and Language Computation -- Human and Machine Language Understandingcited by 1
- 2023Can We Use Probing to Better Understand Fine-tuning and Knowledge Distillation of the BERT NLU?cited by 1
- 2023Rethinking the Construction of Effective Metrics for Understanding the Mechanisms of Pretrained Language Modelscited by 0
- 2025RouterKT: Mixture-of-Experts for Knowledge Tracingcited by 0
- 2026Beyond Decodability: Reconstructing Language Model Representations with an Encoding Probecited by 0
- 2026Beyond Decodability: Reconstructing Language Model Representations with an Encoding Probe1 May 2026
- 2026From Weights to Activations: Is Steering the Next Frontier of Adaptation?15 Apr 2026
- 2025Prophecy: Inferring Formal Properties from Neuron Activations25 Sep 2025
- 2025RouterKT: Mixture-of-Experts for Knowledge Tracing11 Apr 2025
- 2025Representation Engineering for Large-Language Models: Survey and Research Challenges24 Feb 2025
- 2024Multilevel Interpretability Of Artificial Neural Networks: Leveraging Framework And Methods From Neuroscience22 Aug 2024
- 2024Language Models as Models of Language13 Aug 2024
- 2024Vision-Language Models Align with Human Neural Representations in Concept Processing25 Jul 2024
- 2023Rethinking the Construction of Effective Metrics for Understanding the Mechanisms of Pretrained Language Models19 Oct 2023
- 2023Probing Graph Representations7 Mar 2023
- 2023SkinCon: A skin disease dataset densely annotated by domain experts for fine-grained model debugging and analysis1 Feb 2023
- 2023Can We Use Probing to Better Understand Fine-tuning and Knowledge Distillation of the BERT NLU?27 Jan 2023
5 more
- 2023Dissociating language and thought in large language models16 Jan 2023
- 2023Language Cognition and Language Computation -- Human and Machine Language Understanding12 Jan 2023
- 2022Trustworthy Social Bias Measurement20 Dec 2022
- 2021Neural Population Geometry Reveals the Role of Stochasticity in Robust Perception12 Nov 2021
- 2021Conditional probing: measuring usable information beyond a baseline19 Sep 2021
- 2025RouterKT: Mixture-of-Experts for Knowledge Tracingcited by 0
- 2025RouterKT: Mixture-of-Experts for Knowledge Tracing11 Apr 2025
- 2021Conditional probing: measuring usable information beyond a baselinecited by 61
- 2023SkinCon: A skin disease dataset densely annotated by domain experts for fine-grained model debugging and analysiscited by 44
- 2024Language Models as Models of Languagecited by 24
- 2025Representation Engineering for Large-Language Models: Survey and Research Challengescited by 18
- 2021Neural Population Geometry Reveals the Role of Stochasticity in Robust Perceptioncited by 12
- 2023Probing Graph Representationscited by 10
- 2026From Weights to Activations: Is Steering the Next Frontier of Adaptation?cited by 8
- 2022Trustworthy Social Bias Measurementcited by 7
- 2024Multilevel Interpretability Of Artificial Neural Networks: Leveraging Framework And Methods From Neurosciencecited by 5
- 2025Prophecy: Inferring Formal Properties from Neuron Activationscited by 2
- 2023Language Cognition and Language Computation -- Human and Machine Language Understandingcited by 1
- 2023Can We Use Probing to Better Understand Fine-tuning and Knowledge Distillation of the BERT NLU?cited by 1
12 most cited of 13 All 13 (JSON)
- 2026Beyond Decodability: Reconstructing Language Model Representations with an Encoding Probe1 May 2026
- 2026From Weights to Activations: Is Steering the Next Frontier of Adaptation?15 Apr 2026
- 2025Prophecy: Inferring Formal Properties from Neuron Activations25 Sep 2025
- 2025Representation Engineering for Large-Language Models: Survey and Research Challenges24 Feb 2025
- 2024Multilevel Interpretability Of Artificial Neural Networks: Leveraging Framework And Methods From Neuroscience22 Aug 2024
- 2024Language Models as Models of Language13 Aug 2024
- 2023Probing Graph Representations7 Mar 2023
- 2023SkinCon: A skin disease dataset densely annotated by domain experts for fine-grained model debugging and analysis1 Feb 2023
- 2023Can We Use Probing to Better Understand Fine-tuning and Knowledge Distillation of the BERT NLU?27 Jan 2023
- 2023Language Cognition and Language Computation -- Human and Machine Language Understanding12 Jan 2023
- 2022Trustworthy Social Bias Measurement20 Dec 2022
- 2021Neural Population Geometry Reveals the Role of Stochasticity in Robust Perception12 Nov 2021
12 newest of 13 All 13 (JSON)