The Cross-Architecture Substrate: A Domain-Transcendent, Calibration-Surviving Geometric Invariant of Modern Vision Encoders

Different vision neural networks -- trained to classify, contrast, reconstruct, or match images to text -- should have correspondingly different internal representations. We report that they do not. After training, the top sixteen principal directions of variation inside thirteen modern vision encoders converge to the same sixteen-dimensional geometric object. We call this the cross-architecture substrate and study it with PCA, centred kernel alignment (CKA), and Pang 2026 calibration. The substrate transports across four visual domains (natural photographs, medical CT, satellite, microscopy) at median Procrustes-CKA 0.679, and across eight domains (adding sketches, depth, thermal infrared, astronomy) at 0.604, every pair >0.40. It survives Pang calibration globally (7.4x disc-vs-MAE separation, n=13,394) and locally (4.82-5.30, p<10^{-44}). It is not pixel statistics (0.263), not Gabor features (0.31), not a random projection (0.041), and emerges in the first 10% of training while accuracy keeps climbing. We deliver four applications: a label-free transferability filter beating LogME (3x faster, +0.15 Kendall-tau); a four-way domain detector (99.6% accuracy); a frozen low-shot probe (16 dims beat 768-dim DINOv2 by 3.78pp at N=50 labels per class); and a teacher-free distillation auxiliary matching trained-teacher KD on 33 pairs (7.56pp peak gain at 10% label fraction). The substrate does not cross modalities, does not help cross-paradigm distillation, and does not predict transfer quality (rho=0.08 against transfer accuracy).