# How LLMs Follow Instructions: Skillful Coordination, Not a Universal Mechanism

arXiv: https://arxiv.org/abs/2604.06015 · HTML: https://mlsanity.com/paper/2604.06015/

## Key facts

- No citing papers in the map yet (data through 14 Oct 2026).
- Builds on 17 of its 31 references that are in the map.
- First posted on arXiv 7 Apr 2026.

## Builds on (most cited first, 17 of 17)

- [Exploring the Limits of Transfer Learning with a Unified Text-to-Text Transformer](https://mlsanity.com/paper/1910.10683/) (2019)
- [Gemma 2: Improving Open Language Models at a Practical Size](https://mlsanity.com/paper/2408.00118/) (2024)
- [Language Models (Mostly) Know What They Know](https://mlsanity.com/paper/2207.05221/) (2022)
- [Understanding intermediate layers using linear classifier probes](https://mlsanity.com/paper/1610.01644/) (2016)
- [Inference-Time Intervention: Eliciting Truthful Answers from a Language Model](https://mlsanity.com/paper/2306.03341/) (2023)
- [BERT Rediscovers the Classical NLP Pipeline](https://mlsanity.com/paper/1905.05950/) (2019)
- [Probing Classifiers: Promises, Shortcomings, and Advances](https://mlsanity.com/paper/2102.12452/) (2021)
- [Intrinsic Dimensionality Explains the Effectiveness of Language Model Fine-Tuning](https://mlsanity.com/paper/2012.13255/) (2020)
- [Null It Out: Guarding Protected Attributes by Iterative Nullspace Projection](https://mlsanity.com/paper/2004.07667/) (2020)
- [Nuanced Metrics for Measuring Unintended Bias with Real Data for Text Classification](https://mlsanity.com/paper/1903.04561/) (2019)
- [Insights on representational similarity in neural networks with canonical correlation](https://mlsanity.com/paper/1806.05759/) (2018)
- [Improving Instruction-Following in Language Models through Activation Steering](https://mlsanity.com/paper/2410.12877/) (2024)
- [CoEdIT: Text Editing by Task-Specific Instruction Tuning](https://mlsanity.com/paper/2305.09857/) (2023)
- [From Language Modeling to Instruction Following: Understanding the Behavior Shift in LLMs after Instruction Tuning](https://mlsanity.com/paper/2310.00492/) (2023)
- [SAIF: A Sparse Autoencoder Framework for Interpreting and Steering Instruction Following of Language Models](https://mlsanity.com/paper/2502.11356/) (2025)
- [SysBench: Can Large Language Models Follow System Messages?](https://mlsanity.com/paper/2408.10943/) (2024)
- [Do LLMs "know" internally when they follow instructions?](https://mlsanity.com/paper/2410.14516/) (2024)

## Abstract (arXiv)

Instruction tuning is commonly assumed to endow language models with a domain-general ability to follow instructions, yet the underlying mechanism remains poorly understood. Does instruction-following rely on a universal mechanism or compositional skill deployment? We investigate this through diagnostic probing across nine diverse tasks in three instruction-tuned models. Our analysis provides converging evidence against a universal mechanism. First, general probes trained across all tasks show selective rather than uniform deficits relative to task-specific specialists, indicating that representational sharing is partial and structured rather than global. Second, cross-task transfer is weak and clustered by skill similarity. Third, causal ablation reveals sparse asymmetric dependencies rather than shared representations. Tasks also stratify by complexity across layers, with structural constraints emerging early and semantic tasks emerging late. Finally, temporal analysis shows that the constraint signal becomes decodable only once generation is under way, and remains so throughout the response. These findings indicate that instruction-following is better characterized as skillful coordination of diverse linguistic capabilities rather than deployment of a single abstract constraint-checking process.