# The Emergence of Complex Behavior in Large-Scale Ecological Environments

arXiv: https://arxiv.org/abs/2510.18221 · HTML: https://mlsanity.com/paper/2510.18221/

## Key facts

- Cited by 1 papers in the map (data through 14 Oct 2026).
- Builds on 36 of its 77 references that are in the map.
- First posted on arXiv 21 Oct 2025.

## Builds on (most cited first, 30 of 36)

- [Chain-of-Thought Prompting Elicits Reasoning in Large Language Models](https://mlsanity.com/paper/2201.11903/) (2022)
- [Delving Deep into Rectifiers: Surpassing Human-Level Performance on ImageNet Classification](https://mlsanity.com/paper/1502.01852/) (2015)
- [Generative Agents: Interactive Simulacra of Human Behavior](https://mlsanity.com/paper/2304.03442/) (2023)
- [Emergent Abilities of Large Language Models](https://mlsanity.com/paper/2206.07682/) (2022)
- [Evolution Strategies as a Scalable Alternative to Reinforcement Learning](https://mlsanity.com/paper/1703.03864/) (2017)
- [Emergent Tool Use From Multi-Agent Autocurricula](https://mlsanity.com/paper/1909.07528/) (2019)
- [Human-level performance in first-person multiplayer games with population-based deep reinforcement learning](https://mlsanity.com/paper/1807.01281/) (2018)
- [Emergence of Grounded Compositional Language in Multi-Agent Populations](https://mlsanity.com/paper/1703.04908/) (2017)
- [Robots that can adapt like animals](https://mlsanity.com/paper/1407.3501/) (2014)
- [Paired Open-Ended Trailblazer (POET): Endlessly Generating Increasingly Complex and Diverse Learning Environments and Their Solutions](https://mlsanity.com/paper/1901.01753/) (2019)
- [Open-Ended Learning Leads to Generally Capable Agents](https://mlsanity.com/paper/2107.12808/) (2021)
- [Embodied Intelligence via Learning and Evolution](https://mlsanity.com/paper/2102.02202/) (2021)
- [Deep Neuroevolution: Genetic Algorithms Are a Competitive Alternative for Training Deep Neural Networks for Reinforcement Learning](https://mlsanity.com/paper/1712.06567/) (2017)
- [Improving Exploration in Evolution Strategies for Deep Reinforcement Learning via a Population of Novelty-Seeking Agents](https://mlsanity.com/paper/1712.06560/) (2017)
- [Craftax: A Lightning-Fast Benchmark for Open-Ended Reinforcement Learning](https://mlsanity.com/paper/2402.16801/) (2024)
- [Simple random search provides a competitive approach to reinforcement learning](https://mlsanity.com/paper/1803.07055/) (2018)
- [MAgent: A Many-Agent Reinforcement Learning Platform for Artificial Collective Intelligence](https://mlsanity.com/paper/1712.00600/) (2017)
- [Autocurricula and the Emergence of Innovation from Social Interaction: A Manifesto for Multi-Agent Intelligence Research](https://mlsanity.com/paper/1903.00742/) (2019)
- [Reinforcement Learning for Improving Agent Design](https://mlsanity.com/paper/1810.03779/) (2018)
- [Pgx: Hardware-Accelerated Parallel Game Simulators for Reinforcement Learning](https://mlsanity.com/paper/2303.17503/) (2023)
- [Neural MMO: A Massively Multiagent Game Environment for Training and Evaluating Intelligent Agents](https://mlsanity.com/paper/1903.00784/) (2019)
- [Lenia - Biology of Artificial Life](https://mlsanity.com/paper/1812.05433/) (2018)
- [Griddly: A platform for AI research in games](https://mlsanity.com/paper/2011.06363/) (2020)
- [Lenia and Expanded Universe](https://mlsanity.com/paper/2005.03742/) (2020)
- [Megaverse: Simulating Embodied Agents at One Million Experiences per Second](https://mlsanity.com/paper/2107.08170/) (2021)
- [Malthusian Reinforcement Learning](https://mlsanity.com/paper/1812.07019/) (2018)
- [Towards Large-Scale Simulations of Open-Ended Evolution in Continuous Cellular Automata](https://mlsanity.com/paper/2304.05639/) (2023)
- [Biomaker CA: a Biome Maker project using Cellular Automata](https://mlsanity.com/paper/2307.09320/) (2023)
- [Neural MMO 2.0: A Massively Multi-task Addition to Massively Multi-agent Learning](https://mlsanity.com/paper/2311.03736/) (2023)
- [JaxLife: An Open-Ended Agentic Simulator](https://mlsanity.com/paper/2409.00853/) (2024)

## Built on by (most cited first, 1 of 1)

- [Economy of Minds: Emerging Multi-Agent Intelligence with Economic Interactions](https://mlsanity.com/paper/2606.02859/) (2026)

## Abstract (arXiv)

We explore how physical scale and population size shape the emergence of complex behaviors in open-ended ecological environments. In our setting, agents are unsupervised and have no explicit rewards or learning objectives but instead evolve over time according to reproduction, mutation, and selection. As they act, agents also shape their environment and the population around them in an ongoing dynamic ecology. Our goal is not to optimize a single high-performance policy, but instead to examine how behaviors emerge and evolve across large populations due to natural competition and environmental pressures. We use modern hardware along with a new multi-agent simulator to scale the environment and population to sizes much larger than previously attempted, reaching populations of over 60,000 agents, each with their own evolved neural network policy. We identify various emergent behaviors such as long-range resource extraction, vision-based foraging, and predation that arise under competitive and survival pressures. We examine how sensing modalities and environmental scale affect the emergence of these behaviors and find that some of them appear only in sufficiently large environments and populations, and that larger scales increase the stability and consistency of these emergent behaviors. While there is a rich history of research in evolutionary settings, our scaling results on modern hardware provide promising new directions to explore ecology as an instrument of machine learning in an era of increasingly abundant computational resources and efficient machine frameworks. Experimental code is available at https://github.com/jbejjani2022/ecological-emergent-behavior.
