# Neuron-Level Analysis of Cultural Understanding in Large Language Models

arXiv: https://arxiv.org/abs/2510.08284 · HTML: https://mlsanity.com/paper/2510.08284/

## Key facts

- Cited by 1 papers in the map (data through 14 Oct 2026).
- Builds on 30 of its 36 references that are in the map.
- First posted on arXiv 9 Oct 2025.

## Builds on (most cited first, 30 of 30)

- [Attention Is All You Need](https://mlsanity.com/paper/1706.03762/) (2017)
- [BERT: Pre-training of Deep Bidirectional Transformers for Language Understanding](https://mlsanity.com/paper/1810.04805/) (2018)
- [Qwen3 Technical Report](https://mlsanity.com/paper/2505.09388/) (2025)
- [GLUE: A Multi-Task Benchmark and Analysis Platform for Natural Language Understanding](https://mlsanity.com/paper/1804.07461/) (2018)
- [Locating and Editing Factual Associations in GPT](https://mlsanity.com/paper/2202.05262/) (2022)
- [CommonsenseQA: A Question Answering Challenge Targeting Commonsense Knowledge](https://mlsanity.com/paper/1811.00937/) (2018)
- [GLU Variants Improve Transformer](https://mlsanity.com/paper/2002.05202/) (2020)
- [Transformer Feed-Forward Layers Are Key-Value Memories](https://mlsanity.com/paper/2012.14913/) (2020)
- [Mass-Editing Memory in a Transformer](https://mlsanity.com/paper/2210.07229/) (2022)
- [Phi-4 Technical Report](https://mlsanity.com/paper/2412.08905/) (2024)
- [An Empirical Study of Catastrophic Forgetting in Large Language Models During Continual Fine-tuning](https://mlsanity.com/paper/2308.08747/) (2023)
- [Knowledge Neurons in Pretrained Transformers](https://mlsanity.com/paper/2104.08696/) (2021)
- [BLEnD: A Benchmark for LLMs on Everyday Knowledge in Diverse Cultures and Languages](https://mlsanity.com/paper/2406.09948/) (2024)
- [Having Beer after Prayer? Measuring Cultural Bias in Large Language Models](https://mlsanity.com/paper/2305.14456/) (2023)
- [Language-Specific Neurons: The Key to Multilingual Capabilities in Large Language Models](https://mlsanity.com/paper/2402.16438/) (2024)
- [On the Multilingual Ability of Decoder-based Pre-trained Language Models: Finding and Controlling Language-Specific Neurons](https://mlsanity.com/paper/2404.02431/) (2024)
- [The Semantic Hub Hypothesis: Language Models Share Semantic Representations Across Languages and Modalities](https://mlsanity.com/paper/2411.04986/) (2024)
- [WorldValuesBench: A Large-Scale Benchmark Dataset for Multi-Cultural Value Awareness of Language Models](https://mlsanity.com/paper/2404.16308/) (2024)
- [Lost in Multilinguality: Dissecting Cross-lingual Factual Inconsistency in Transformer Language Models](https://mlsanity.com/paper/2504.04264/) (2025)
- [Competition of Mechanisms: Tracing How Language Models Handle Facts and Counterfactuals](https://mlsanity.com/paper/2402.11655/) (2024)
- [Neuron-Level Knowledge Attribution in Large Language Models](https://mlsanity.com/paper/2312.12141/) (2023)
- [Self-Pluralising Culture Alignment for Large Language Models](https://mlsanity.com/paper/2410.12971/) (2024)
- [An Evaluation of Cultural Value Alignment in LLM](https://mlsanity.com/paper/2504.08863/) (2025)
- [Mitigating Biases for Instruction-following Language Models via Bias Neurons Elimination](https://mlsanity.com/paper/2311.09627/) (2023)
- [Cultural Learning-Based Culture Adaptation of Language Models](https://mlsanity.com/paper/2504.02953/) (2025)
- [Disentangling Language and Culture for Evaluating Multilingual Large Language Models](https://mlsanity.com/paper/2505.24635/) (2025)
- [Identifying Query-Relevant Neurons in Large Language Models for Long-Form Texts](https://mlsanity.com/paper/2406.10868/) (2024)
- [Unveiling the Lexical Sensitivity of LLMs: Combinatorial Optimization for Prompt Enhancement](https://mlsanity.com/paper/2405.20701/) (2024)
- [Neuron-based Personality Trait Induction in Large Language Models](https://mlsanity.com/paper/2410.12327/) (2024)
- [Isolating Culture Neurons in Multilingual Large Language Models](https://mlsanity.com/paper/2508.02241/) (2025)

## Built on by (most cited first, 1 of 1)

- [Cultural Binding Heads in Language Models](https://mlsanity.com/paper/2605.28543/) (2026)

## Abstract (arXiv)

As large language models (LLMs) are increasingly deployed worldwide, ensuring their fair and comprehensive cultural understanding is important. However, LLMs exhibit cultural bias and limited awareness of underrepresented cultures, while the mechanisms underlying their cultural understanding remain underexplored. To fill this gap, we conduct a neuron-level analysis to identify neurons that drive cultural behavior, introducing a gradient-based scoring method with additional filtering for precise refinement. We identify culture-general neurons contributing to cultural understanding regardless of cultures, and culture-specific neurons tied to an individual culture. Culture-general and culture-specific neurons account for less than 1% of all neurons and are concentrated in shallow to middle MLP layers. We validate their role by showing that suppressing them substantially degrades performance on cultural benchmarks (by up to 30%), while performance on general natural language understanding (NLU) benchmarks remains largely unaffected. Moreover, we show that culture-specific neurons support knowledge of not only the target culture, but also related cultures. Finally, we demonstrate that training on NLU benchmarks can diminish models' cultural understanding when we update modules containing many culture-general neurons. These findings provide insights into the internal mechanisms of LLMs and offer practical guidance for model training and engineering. Our code is available at https://github.com/ynklab/CULNIG