# Discovering Influential Neuron Path in Vision Transformers

arXiv: https://arxiv.org/abs/2503.09046 · HTML: https://mlsanity.com/paper/2503.09046/

## Key facts

- Cited by 7 papers in the map (data through 14 Oct 2026).
- Builds on 45 of its 51 references that are in the map.
- First posted on arXiv 12 Mar 2025.

## Builds on (most cited first, 30 of 45)

- [Attention Is All You Need](https://mlsanity.com/paper/1706.03762/) (2017)
- [BERT: Pre-training of Deep Bidirectional Transformers for Language Understanding](https://mlsanity.com/paper/1810.04805/) (2018)
- [Learning Transferable Visual Models From Natural Language Supervision](https://mlsanity.com/paper/2103.00020/) (2021)
- [An Image is Worth 16x16 Words: Transformers for Image Recognition at Scale](https://mlsanity.com/paper/2010.11929/) (2020)
- [DINOv2: Learning Robust Visual Features without Supervision](https://mlsanity.com/paper/2304.07193/) (2023)
- [Masked Autoencoders Are Scalable Vision Learners](https://mlsanity.com/paper/2111.06377/) (2021)
- [Emerging Properties in Self-Supervised Vision Transformers](https://mlsanity.com/paper/2104.14294/) (2021)
- [Grad-CAM: Visual Explanations from Deep Networks via Gradient-based Localization](https://mlsanity.com/paper/1610.02391/) (2016)
- [Scalable Diffusion Models with Transformers](https://mlsanity.com/paper/2212.09748/) (2022)
- [Axiomatic Attribution for Deep Networks](https://mlsanity.com/paper/1703.01365/) (2017)
- [Learning Deep Features for Discriminative Localization](https://mlsanity.com/paper/1512.04150/) (2015)
- [Visualizing and Understanding Convolutional Networks](https://mlsanity.com/paper/1311.2901/) (2013)
- [Understanding Black-box Predictions via Influence Functions](https://mlsanity.com/paper/1703.04730/) (2017)
- [Transformer Feed-Forward Layers Are Key-Value Memories](https://mlsanity.com/paper/2012.14913/) (2020)
- [Interpretability Beyond Feature Attribution: Quantitative Testing with Concept Activation Vectors (TCAV)](https://mlsanity.com/paper/1711.11279/) (2017)
- [SmoothGrad: removing noise by adding noise](https://mlsanity.com/paper/1706.03825/) (2017)
- [Grad-CAM++: Improved Visual Explanations for Deep Convolutional Networks](https://mlsanity.com/paper/1710.11063/) (2017)
- [Network Dissection: Quantifying Interpretability of Deep Visual Representations](https://mlsanity.com/paper/1704.05796/) (2017)
- [Transformer Interpretability Beyond Attention Visualization](https://mlsanity.com/paper/2012.09838/) (2020)
- [Knowledge Neurons in Pretrained Transformers](https://mlsanity.com/paper/2104.08696/) (2021)
- [Fast Model Editing at Scale](https://mlsanity.com/paper/2110.11309/) (2021)
- [Intriguing Properties of Vision Transformers](https://mlsanity.com/paper/2105.10497/) (2021)
- [A Multiscale Visualization of Attention in the Transformer Model](https://mlsanity.com/paper/1906.05714/) (2019)
- [On Explainability of Graph Neural Networks via Subgraph Explorations](https://mlsanity.com/paper/2102.05152/) (2021)
- [Mixture-of-Depths: Dynamically allocating compute in transformer-based language models](https://mlsanity.com/paper/2404.02258/) (2024)
- [Vision Transformers are Robust Learners](https://mlsanity.com/paper/2105.07581/) (2021)
- [Visualizing and Understanding Neural Models in NLP](https://mlsanity.com/paper/1506.01066/) (2015)
- [Visualizing and Understanding Recurrent Networks](https://mlsanity.com/paper/1506.02078/) (2015)
- [CLIP-Dissect: Automatic Description of Neuron Representations in Deep Vision Networks](https://mlsanity.com/paper/2204.10965/) (2022)
- [Natural Language Descriptions of Deep Visual Features](https://mlsanity.com/paper/2201.11114/) (2022)

## Built on by (most cited first, 7 of 7)

- [TraceRouter: Robust Safety for Large Foundation Models via Path-Level Intervention](https://mlsanity.com/paper/2601.21900/) (2026)
- [Why LVLMs Are More Prone to Hallucinations in Longer Responses: The Role of Context](https://mlsanity.com/paper/2510.20229/) (2025)
- [Cross-Modal Unlearning via Influential Neuron Path Editing in Multimodal Large Language Models](https://mlsanity.com/paper/2511.06793/) (2025)
- [Granular Concept Circuits: Toward a Fine-Grained Circuit Discovery for Concept Representations](https://mlsanity.com/paper/2508.01728/) (2025)
- [Demystifying Variance in Circuit Discovery of LLMs](https://mlsanity.com/paper/2606.16920/) (2026)
- [Seeing Through Circuits: Faithful Mechanistic Interpretability for Vision Transformers](https://mlsanity.com/paper/2604.14477/) (2026)
- [Are the High-weight Neurons the Important Ones in Image Classification Neural Networks?](https://mlsanity.com/paper/2607.25529/) (2026)

## Abstract (arXiv)

Vision Transformer models exhibit immense power yet remain opaque to human understanding, posing challenges and risks for practical applications. While prior research has attempted to demystify these models through input attribution and neuron role analysis, there's been a notable gap in considering layer-level information and the holistic path of information flow across layers. In this paper, we investigate the significance of influential neuron paths within vision Transformers, which is a path of neurons from the model input to output that impacts the model inference most significantly. We first propose a joint influence measure to assess the contribution of a set of neurons to the model outcome. And we further provide a layer-progressive neuron locating approach that efficiently selects the most influential neuron at each layer trying to discover the crucial neuron path from input to output within the target model. Our experiments demonstrate the superiority of our method finding the most influential neuron path along which the information flows, over the existing baseline solutions. Additionally, the neuron paths have illustrated that vision Transformers exhibit some specific inner working mechanism for processing the visual information within the same image category. We further analyze the key effects of these neurons on the image classification task, showcasing that the found neuron paths have already preserved the model capability on downstream tasks, which may also shed some lights on real-world applications like model pruning. The project website including implementation code is available at https://foundation-model-research.github.io/NeuronPath/.