# Finding Neurons in a Haystack: Case Studies with Sparse Probing

arXiv: https://arxiv.org/abs/2305.01610 · HTML: https://mlsanity.com/paper/2305.01610/

## Key facts

- Cited by 334 papers in the map (data through 14 Oct 2026).
- Builds on 71 of its 107 references that are in the map.
- First posted on arXiv 2 May 2023.

## Builds on (most cited first, 30 of 71)

- [Language Models are Few-Shot Learners](https://mlsanity.com/paper/2005.14165/) (2020)
- [Layer Normalization](https://mlsanity.com/paper/1607.06450/) (2016)
- [PaLM: Scaling Language Modeling with Pathways](https://mlsanity.com/paper/2204.02311/) (2022)
- [Gaussian Error Linear Units (GELUs)](https://mlsanity.com/paper/1606.08415/) (2016)
- [Locating and Editing Factual Associations in GPT](https://mlsanity.com/paper/2202.05262/) (2022)
- [Sparks of Artificial General Intelligence: Early experiments with GPT-4](https://mlsanity.com/paper/2303.12712/) (2023)
- [Emergent Abilities of Large Language Models](https://mlsanity.com/paper/2206.07682/) (2022)
- [The Pile: An 800GB Dataset of Diverse Text for Language Modeling](https://mlsanity.com/paper/2101.00027/) (2020)
- [Pythia: A Suite for Analyzing Large Language Models Across Training and Scaling](https://mlsanity.com/paper/2304.01373/) (2023)
- [Understanding intermediate layers using linear classifier probes](https://mlsanity.com/paper/1610.01644/) (2016)
- [Transformer Feed-Forward Layers Are Key-Value Memories](https://mlsanity.com/paper/2012.14913/) (2020)
- [Interpretability Beyond Feature Attribution: Quantitative Testing with Concept Activation Vectors (TCAV)](https://mlsanity.com/paper/1711.11279/) (2017)
- [Interpretability in the Wild: a Circuit for Indirect Object Identification in GPT-2 small](https://mlsanity.com/paper/2211.00593/) (2022)
- [Sanity Checks for Saliency Maps](https://mlsanity.com/paper/1810.03292/) (2018)
- [BERT Rediscovers the Classical NLP Pipeline](https://mlsanity.com/paper/1905.05950/) (2019)
- [Toy Models of Superposition](https://mlsanity.com/paper/2209.10652/) (2022)
- [In-context Learning and Induction Heads](https://mlsanity.com/paper/2209.11895/) (2022)
- [Probing Classifiers: Promises, Shortcomings, and Advances](https://mlsanity.com/paper/2102.12452/) (2021)
- [InfoGAN: Interpretable Representation Learning by Information Maximizing Generative Adversarial Nets](https://mlsanity.com/paper/1606.03657/) (2016)
- [Progress measures for grokking via mechanistic interpretability](https://mlsanity.com/paper/2301.05217/) (2023)
- [A Primer in BERTology: What we know about how BERT works](https://mlsanity.com/paper/2002.12327/) (2020)
- [Discovering Latent Knowledge in Language Models Without Supervision](https://mlsanity.com/paper/2212.03827/) (2022)
- [Network Dissection: Quantifying Interpretability of Deep Visual Representations](https://mlsanity.com/paper/1704.05796/) (2017)
- [Designing and Interpreting Probes with Control Tasks](https://mlsanity.com/paper/1909.03368/) (2019)
- [Disentangling by Factorising](https://mlsanity.com/paper/1802.05983/) (2018)
- [Transformer Feed-Forward Layers Build Predictions by Promoting Concepts in the Vocabulary Space](https://mlsanity.com/paper/2203.14680/) (2022)
- [What you can cram into a single vector: Probing sentence embeddings for linguistic properties](https://mlsanity.com/paper/1805.01070/) (2018)
- [What do you learn from context? Probing for sentence structure in contextualized word representations](https://mlsanity.com/paper/1905.06316/) (2019)
- [Emergent World Representations: Exploring a Sequence Model Trained on a Synthetic Task](https://mlsanity.com/paper/2210.13382/) (2022)
- [DeCAF: A Deep Convolutional Activation Feature for Generic Visual Recognition](https://mlsanity.com/paper/1310.1531/) (2013)

## Built on by (most cited first, 30 of 334)

- [The Geometry of Truth: Emergent Linear Structure in Large Language Model Representations of True/False Datasets](https://mlsanity.com/paper/2310.06824/) (2023)
- [Scaling and evaluating sparse autoencoders](https://mlsanity.com/paper/2406.04093/) (2024)
- [Mechanistic Interpretability for AI Safety -- A Review](https://mlsanity.com/paper/2404.14082/) (2024)
- [Gemma Scope: Open Sparse Autoencoders Everywhere All At Once on Gemma 2](https://mlsanity.com/paper/2408.05147/) (2024)
- [Towards Best Practices of Activation Patching in Language Models: Metrics and Methods](https://mlsanity.com/paper/2309.16042/) (2023)
- [Emergent Linear Representations in World Models of Self-Supervised Sequence Models](https://mlsanity.com/paper/2309.00941/) (2023)
- [Language Models Represent Space and Time](https://mlsanity.com/paper/2310.02207/) (2023)
- [Jumping Ahead: Improving Reconstruction Fidelity with JumpReLU Sparse Autoencoders](https://mlsanity.com/paper/2407.14435/) (2024)
- [Do Llamas Work in English? On the Latent Language of Multilingual Transformers](https://mlsanity.com/paper/2402.10588/) (2024)
- [The Unreasonable Ineffectiveness of the Deeper Layers](https://mlsanity.com/paper/2403.17887/) (2024)
- [Transcoders Find Interpretable LLM Feature Circuits](https://mlsanity.com/paper/2406.11944/) (2024)
- [Improving Dictionary Learning with Gated Sparse Autoencoders](https://mlsanity.com/paper/2404.16014/) (2024)
- [CROMA: Remote Sensing Representations with Contrastive Radar-Optical Masked Autoencoders](https://mlsanity.com/paper/2311.00566/) (2023)
- [The pitfalls of next-token prediction](https://mlsanity.com/paper/2403.06963/) (2024)
- [A is for Absorption: Studying Feature Splitting and Absorption in Sparse Autoencoders](https://mlsanity.com/paper/2409.14507/) (2024)
- [The Geometry of Categorical and Hierarchical Concepts in Large Language Models](https://mlsanity.com/paper/2406.01506/) (2024)
- [The Clock and the Pizza: Two Stories in Mechanistic Explanation of Neural Networks](https://mlsanity.com/paper/2306.17844/) (2023)
- [The Hydra Effect: Emergent Self-repair in Language Model Computations](https://mlsanity.com/paper/2307.15771/) (2023)
- [Does Circuit Analysis Interpretability Scale? Evidence from Multiple Choice Capabilities in Chinchilla](https://mlsanity.com/paper/2307.09458/) (2023)
- [The Remarkable Robustness of LLMs: Stages of Inference?](https://mlsanity.com/paper/2406.19384/) (2024)
- [Llama Scope: Extracting Millions of Features from Llama-3.1-8B with Sparse Autoencoders](https://mlsanity.com/paper/2410.20526/) (2024)
- [Language-Specific Neurons: The Key to Multilingual Capabilities in Large Language Models](https://mlsanity.com/paper/2402.16438/) (2024)
- [SAEBench: A Comprehensive Benchmark for Sparse Autoencoders in Language Model Interpretability](https://mlsanity.com/paper/2503.09532/) (2025)
- [Future Lens: Anticipating Subsequent Tokens from a Single Hidden State](https://mlsanity.com/paper/2311.04897/) (2023)
- [A Survey on Uncertainty Quantification of Large Language Models: Taxonomy, Open Research Challenges, and Future Directions](https://mlsanity.com/paper/2412.05563/) (2024)
- [A Practical Review of Mechanistic Interpretability for Transformer-Based Language Models](https://mlsanity.com/paper/2407.02646/) (2024)
- [Are Sparse Autoencoders Useful? A Case Study in Sparse Probing](https://mlsanity.com/paper/2502.16681/) (2025)
- [Automatically Interpreting Millions of Features in Large Language Models](https://mlsanity.com/paper/2410.13928/) (2024)
- [Rethinking Interpretability in the Era of Large Language Models](https://mlsanity.com/paper/2402.01761/) (2024)
- [AtP*: An efficient and scalable method for localizing LLM behaviour to components](https://mlsanity.com/paper/2403.00745/) (2024)

## Abstract (arXiv)

Despite rapid adoption and deployment of large language models (LLMs), the internal computations of these models remain opaque and poorly understood. In this work, we seek to understand how high-level human-interpretable features are represented within the internal neuron activations of LLMs. We train k-sparse linear classifiers (probes) on these internal activations to predict the presence of features in the input; by varying the value of k we study the sparsity of learned representations and how this varies with model scale. With k=1, we localize individual neurons which are highly relevant for a particular feature, and perform a number of case studies to illustrate general properties of LLMs. In particular, we show that early layers make use of sparse combinations of neurons to represent many features in superposition, that middle layers have seemingly dedicated neurons to represent higher-level contextual features, and that increasing scale causes representational sparsity to increase on average, but there are multiple types of scaling dynamics. In all, we probe for over 100 unique features comprising 10 different categories in 7 different models spanning 70 million to 6.9 billion parameters.